[1]郑绿珠,李晓阳,蔡志全.植物激素调控色素合成的分子机制研究进展[J].江苏农业学报,2026,42(06):1270-1282.[doi:doi:10.3969/j.issn.1000-4440.2026.06.019]
　ZHENG Lüzhu,LI Xiaoyang,CAI Zhiquan.Research progress on the molecular mechanisms of plant hormones in pigment synthesis[J].,2026,42(06):1270-1282.[doi:doi:10.3969/j.issn.1000-4440.2026.06.019]
点击复制

植物激素调控色素合成的分子机制研究进展()

江苏农业学报[ISSN:1006-6977/CN:61-1281/TN]

卷:
42
期数:
2026年06期
页码:
1270-1282
栏目:
综述
出版日期:
2026-06-30

文章信息/Info

Title:
Research progress on the molecular mechanisms of plant hormones in pigment synthesis
作者:
郑绿珠; 李晓阳; 蔡志全
(佛山大学农业与生物工程学院,广东佛山528231)
Author(s):
ZHENG Lüzhu; LI Xiaoyang; CAI Zhiquan
(School of Agricultural and Biological Engineering, Foshan University, Foshan 528231, China)
关键词:
色素代谢; 植物激素; 转录调控; 园艺植物; 分子育种
Keywords:
pigment metabolism; plant hormone; transcriptional regulation; horticultural plants; molecular breeding
分类号:
S946.885
DOI:
doi:10.3969/j.issn.1000-4440.2026.06.019
文献标志码:
A
摘要:
植物色素(花青素、类胡萝卜素和叶绿素)是评价园艺作物观赏价值和经济价值的重要指标,园艺作物生长发育过程中主要色素物质合成与积累的变化受到自身遗传背景、发育阶段以及环境的调控。此外,植物激素在园艺作物色素代谢中发挥重要作用,色素的合成和降解均受到内源激素变化和外源激素处理的影响而发生变化。本文通过综述植物激素（脱落酸、生长素、乙烯、茉莉酸、褪黑素、油菜素内酯）在园艺作物色素合成与积累中的作用、激素互作以及植物激素通过转录因子的调控影响园艺作物的色素合成与积累等多方面的研究进展,以期为园艺作物分子育种提供参考。
Abstract:
Plant pigments (anthocyanins, carotenoids, and chlorophyll) are important indicators of the ornamental and economic value of horticultural crops. The changes in the synthesis and accumulation of these primary pigments during the growth and development of horticultural crops are regulated by their genetic background, developmental stages, and environmental conditions. Additionally, plant hormones play a crucial role in the metabolism of pigments in horticultural crops, as both the synthesis and degradation of pigments are influenced by changes in endogenous hormones and treatments with exogenous hormones. This paper reviewed the roles of plant hormones such as abscisic acid, auxins, ethylene, jasmo-nic acid, melatonin, and brassinosteroids in the synthesis and accumulation of pigments in horticultural crops, the interactions between hormones, and how plant hormones regulate pigment synthesis and accumulation in horticultural crops through transcription factors. The aim is to provide reference directions for molecular breeding of horticultural crops.

参考文献/References:

[1]ZOU J Q, HUANG S N, GAO Y, et al. Mutation in BrFLS encoding flavonol synthase induced anthocyanin accumulation in Chinese cabbage[J]. Theoretical and Applied Genetics,2024,137(2):44.
[2]TIAN J, HAN Z Y, ZHANG J, et al. The balance of expression of dihydroflavonol 4-reductase and flavonol synthase regulates flavonoid biosynthesis and red foliage coloration in crabapples[J]. Scientific Reports,2015,5:12228.
[3]LI J Y, SUN J L, TIAN Y Y, et al. The FtsH-inactive protein FtsHi5 is required for chloroplast development and protein accumulation in chloroplasts at low ambient temperature in Arabidopsis[J]. Frontiers in Plant Science,2022,12:830390.
[4]CAI X, YANG J, ZENG X L, et al. Light and temperature effects on the color and scent of Osmanthus fragrans ‘houban Yingui’[J]. Horticultural Science and Technology,2022,40(6):595-604.
[5]ZHANG Y F, LIU Y, LING L, et al. Phenotypic,physiological,and molecular response of Loropetalum chinense var.rubrum under different light quality treatments based on leaf color changes[J]. Plants,2023,12(11):2169.
[6]LI X B, ZHANG D D, PAN X H, et al. Regulation of carotenoid metabolism and ABA biosynthesis during blueberry fruit ripening[J]. Plant Physiology and Biochemistry,2024,206:108232.
[7]FENG S Y, LIU Z, CHEN H L, et al. PHGD an integrative and user-friendly database for plant hormone-related genes[J]. iMeta,2024,3(1):e164.
[8]YOSHIDA T, FERNIE A R. Hormonal regulation of plant primary metabolism under drought[J]. Journal of Experimental Botany,2024,75(6):1714-1725.
[9]LIU Y C, DONG B, ZHANG C, et al. Effects of exogenous abscisic acid (ABA) on carotenoids and petal color in Osmanthus fragrans ‘yanhonggui’[J]. Plants,2020,9(4):454.
[10]邓昌哲,秦于玲,李开绵,等. 外源ABA对木薯叶片β-胡萝卜素合成通路相关基因表达的影响[J]. 热带作物学报,2017,38(4):667-672.
[11]ACEVEDO O, PONCE C, ARELLANO M, et al. ABA biosynthesis- and signaling-related gene expression differences between sweet cherry fruits suggest attenuation of ABA pathway in bicolored cultivars[J]. Plants,2023,12(13):2493.
[12]GUO S H, ZHANG M, FENG M X, et al. miR156b-targeted VvSBP8/13 functions downstream of the abscisic acid signal to regulate anthocyanins biosynthesis in grapevine fruit under drought[J]. Horticulture Research,2024,11(2):uhad293.
[13]LI J J, GAO Q F, QUAN Y, et al. VvERF045 integrates ethylene,brassinosteroid and abscisic acid pathways to orchestrate anthocyanin biosynthesis in grapevine[J]. Plant Biotechnology Journal,2026,24(2):787-809.
[14]LEI K J, ZHOU H, GU D L, et al. The involvement of abscisic acid-insensitive mutants in low phosphate stress responses during rhizosphere acidification,anthocyanin accumulation and Pi homeostasis in Arabidopsis[J]. Plant Science,2022,322:111358.
[15]MITALO O W, OTSUKI T, OKADA R, et al. Low temperature modulates natural peel degreening in lemon fruit independently of endogenous ethylene[J]. Journal of Experimental Botany,2020,71(16):4778-4796.
[16]CHEN R C, MAO L C, GUAN W L, et al. ABA-mediated miR5290 promotes anthocyanin biosynthesis by inhibiting the expression of FaMADS1 in postharvest strawberry fruit[J]. Postharvest Biology and Technology,2022,189:111934.
[17]李钰卓. 菜心BrNAP基因的克隆及其在采后叶片衰老中的功能研究[D]. 杭州:浙江大学,2020.
[18]SUN Q, HE Z C, FENG D, et al. The abscisic acid-responsive transcriptional regulatory module CsERF110-CsERF53 orchestrates Citrus fruit coloration[J]. Plant Communications,2024,5(11):101065.
[19]WANG Y C, WANG N, XU H F, et al. Auxin regulates anthocyanin biosynthesis through the Aux/IAA-ARF signaling pathway in apple[J]. Horticulture Research,2018,5:59.
[20]WANG C K, HAN P L, ZHAO Y W, et al. Auxin regulates anthocyanin biosynthesis through the auxin repressor protein MdIAA26[J]. Biochemical and Biophysical Research Communications,2020,533(4):717-722.
[21]MA X, LI D, CAI J, et al. Auxin response factor, RhARF8 contributes to rose flower color fading via regulating anthocyanin biosynthesis by directly activating RhCHSa/c promoter activity[J]. Ornamental Plant Research,2025,5(1):1-12.
[22]CLAYTON-CUCH D, YU L, SHIRLEY N, et al. Auxin treatment enhances anthocyanin production in the non-climacteric sweet cherry (Prunus avium L.)[J]. International Journal of Molecular Sciences,2021,22(19):10760.
[23]FENG S Y, LI N, CHEN H L, et al. Large-scale analysis of the ARF and Aux/IAA gene families in 406 horticultural and other plants[J]. Molecular Horticulture,2024,4(1):13.
[24]ZHANG D Y, WU S D, LI N, et al. Chemical induction of leaf senescence and powdery mildew resistance involves ethylene-mediated chlorophyll degradation and ROS metabolism in cucumber[J]. Horticulture Research,2022,9:uhac101.
[25]CHEN H C, LAI X H, WANG L H, et al. Ethylene response factor MaERF012 modulates fruit ripening by regulating chlorophyll degradation and softening in banana[J]. Foods,2022,11(23):3882.
[26]曹声海.响应乙烯的转录因子在月季花朵衰老过程中的功能分析[D]. 武汉:华中农业大学,2016.
[27]DARVISH M, SHIRZAD H, ASGHARI M, et al. 24-epibrasinolide modulates the vase life of Lisianthus cut flowers by modulating ACC oxidase enzyme activity and physiological responses[J]. Plants,2021,10(5):1024.
[28]SUN Q Q, LIU L, ZHANG L, et al. Melatonin promotes carotenoid biosynthesis in an ethylene-dependent manner in tomato fruits[J]. Plant Science,2020,298:110580.
[29]WEI Y, JIN J T, XU Y X, et al. Ethylene-activated MdPUB24 mediates ubiquitination of MdBEL7 to promote chlorophyll degradation in apple fruit[J]. The Plant Journal,2021,108(1):169-182.
[30]FOO E, ROSS J J, DAVIES N W, et al. A role for ethylene in the phytochrome-mediated control of vegetative development[J]. The Plant Journal,2006,46(6):911-921.
[31]CAO Y W, SONG M, BI M M, et al. Lily (Lilium spp.) LhERF4 negatively affects anthocyanin biosynthesis by suppressing LhMYBSPLATTER transcription[J]. Plant Science,2024,342:112026.
[32]ZHANG Y B, SUN Y B, DU W F, et al. Ethylene promotes anthocyanin synthesis in ‘Viviana’ lily via the LvMYB5-LvERF113-LvMYB1 module[J]. Horticulture Research,2025,12(6):uhaf059.
[33]ZHANG H D, ZHANG K P, ZHAO X Y, et al. Galactinol synthase 2 influences the metabolism of chlorophyll,carotenoids,and ethylene in tomato fruits[J]. Journal of Experimental Botany,2024,75(11):3337-3350.
[34]KHUNMUANG S, KANLAYANARAT S, WONGS-AREE C, et al. Ethylene induces a rapid degradation of petal anthocyanins in cut Vanda ‘Sansai blue’ orchid flowers[J]. Frontiers in Plant Science,2019,10:1004.
[35]JEONG S W, DAS P K, JEOUNG S C, et al. Ethylene suppression of sugar-induced anthocyanin pigmentation in Arabidopsis[J]. Plant Physiology,2010,154(3):1514-1531.
[36]GUO P, HUANG Z Q, ZHAO W, et al. Mechanisms for leaf color changes in Osmanthus fragrans ‘Ziyan Gongzhu’ using physiology,transcriptomics and metabolomics[J]. BMC Plant Biology,2023,23(1):453.
[37]NI J B, WANG S M, YU W J, et al. The ethylene-responsive transcription factor PpERF9 represses PpRAP2.4 and PpMYB114 via histone deacetylation to inhibit anthocyanin biosynthesis in pear[J]. The Plant Cell,2023,35(6):2271-2292.
[38]WANG S, LI L X, FANG Y, et al. MdERF1B-MdMYC2 module integrates ethylene and jasmonic acid to regulate the biosynthesis of anthocyanin in apple[J]. Horticulture Research,2022,9:uhac142.
[39]LIU S H, FANG S, LIU C L, et al. Transcriptomics integrated with metabolomics reveal the effects of ultraviolet-B radiation on flavonoid biosynthesis in Antarctic moss[J]. Frontiers in Plant Science,2021,12:788377.
[40]LUO X Y, LI Z H, XIAO S H, et al. Phosphate deficiency enhances cotton resistance to Verticillium dahliae through activating jasmonic acid biosynthesis and phenylpropanoid pathway[J]. Plant Science,2021,302:110724.
[41]QIU X, XU Y H, XIONG B, et al. Effects of exogenous methyl jasmonate on the synthesis of endogenous jasmonates and the regulation of photosynthesis in Citrus[J]. Physiologia Plantarum,2020,170(3):398-414.
[42]WANG H H, WANG X Q, YU C Y, et al. MYB transcription factor PdMYB118 directly interacts with bHLH transcription factor PdTT8 to regulate wound-induced anthocyanin biosynthesis in poplar[J]. BMC Plant Biology,2020,20(1):173.
[43]LI T, JIA K P, LIAN H L, et al. Jasmonic acid enhancement of anthocyanin accumulation is dependent on phytochrome A signaling pathway under far-red light in Arabidopsis[J]. Biochemical and Biophysical Research Communications,2014,454(1):78-83.
[44]ULLAH A, AKBAR A, YANG X Y. Jasmonic acid (JA)-mediated signaling in leaf senescence[M]//Senescence Signalling and Control in Plants.Amsterdam:Elsevier,2019:111-123.
[45]JI X L, ZHAO L L, LIU B Y, et al. MdZFP7 integrates JA and GA signals via interaction with MdJAZ2 and MdRGL3a in regulating anthocyanin biosynthesis and undergoes degradation by the E3 ubiquitin ligase MdBRG3[J]. Journal of Integrative Plant Biology,2025,67(5):1339-1363.
[46]LI H L, XU R R, GUO X L, et al. The MdNAC72-MdABI5 module acts as an interface integrating jasmonic acid and gibberellin signals and undergoes ubiquitination-dependent degradation regulated by MdSINA2 in apple[J]. New Phytologist,2024,243(3):997-1016.
[47]LI X B, NIU G Q, FAN Y H, et al. Synthetic dual hormone-responsive promoters enable engineering of plants with broad-spectrum resistance[J]. Plant Communications,2023,4(4):100596.
[48]WANG Y, WANG H, XIN Y, et al. Genome-wide identification of SmJAZ gene family in eggplant and functional mechanism analysis of SmJAZ9 modulating high temperature and darkness stress-regulated anthocyanin biosynthesis[J]. Vegetable Research,2025,5(1):1-14.
[49]SU Y N, HUANG Y Z, DONG X T, et al. Exogenous methyl jasmonate improves heat tolerance of perennial ryegrass through alteration of osmotic adjustment,antioxidant defense,and expression of jasmonic acid-responsive genes[J]. Frontiers in Plant Science,2021,12:664519.
[50]罗浩. 外源茉莉酸甲酯调控发芽玉米籽粒合成类胡萝卜素的研究[D]. 哈尔滨:东北林业大学,2020.
[51]ZENG W, MOSTAFA S, LU Z G, et al. Melatonin-mediated abiotic stress tolerance in plants[J]. Frontiers in Plant Science,2022,13:847175.
[52]PAN Y, XU X S, LI L, et al. Melatonin-mediated development and abiotic stress tolerance in plants[J]. Frontiers in Plant Science,2023,14:1100827.
[53]COLOMBAGE R, SINGH M B, BHALLA P L. Melatonin and abiotic stress tolerance in crop plants[J]. International Journal of Molecular Sciences,2023,24(8):7447.
[54]AGHDAM M S, LUO Z S, LI L, et al. Melatonin treatment maintains nutraceutical properties of pomegranate fruits during cold storage[J]. Food Chemistry,2020,303:125385.
[55]周光怡,刘晓娟,王睿,等. 外源褪黑素对高粱苗期冷害的缓解作用研究[J]. 山东农业科学,2025,57(2):55-63.
[56]梁佳,胡朝阳,谢志明,等. 外源褪黑素缓解甜高粱幼苗干旱胁迫的生理效应[J]. 草业学报,2023,32(7):206-215.
[57]SUN H L, WANG X Y, GU G F, et al. The PuERF27-PuMYB10-PuGSTF12 module regulates melatonin-induced anthocyanin biosynthesis in pear fruits[J]. Journal of Agricultural and Food Chemistry,2025,73(19):11780-11792.
[58]CHEN L, TIAN J, WANG S F, et al. Application of melatonin promotes anthocyanin accumulation in crabapple leaves[J]. Plant Physiology and Biochemistry,2019,142:332-341.
[59]杨艳,肖斌. 干旱胁迫下外源褪黑素对王族海棠光合作用、ASA-GSH循环以及激素变化的影响[J]. 河南农业科学,2024,53(6):100-110.
[60]蒋朝维,吴俊杰,姜浩,等. 低温胁迫下烤烟幼苗对喷施外源褪黑素的生理响应[J]. 江苏农业科学,2023,51(21):62-66.
[61]LI J W, YANG J, LIANG X， et al. Melatonin-mediated regulation of antioxidant defense enhances the resistance of tea plants (Camellia sinensis L.) to lead-induced stress[J]. Plants,2025,14(10):1520.
[62]LIANG D, SHEN Y Q, NI Z Y, et al. Exogenous melatonin application delays senescence of kiwifruit leaves by regulating the antioxidant capacity and biosynthesis of flavonoids[J]. Frontiers in Plant Science,2018,9:426.
[63]张玉萍,刘威生,孙绍春,等. 油菜素内酯在果树上的应用研究进展[J]. 安徽农业科学,2010,38(12):6153-6154,6157.
[64]MU D W, FENG N J, ZHENG D F, et al. Physiological mechanism of exogenous brassinolide alleviating salt stress injury in rice seedlings[J]. Scientific Reports,2022,12:20439.
[65]靳开川,何金环. 油菜素内酯在植物抗逆中的作用及信号传导机制综述[J]. 江苏农业科学,2017,45(14):4-7.
[66]陈晨,程大伟,李兰,等. 油菜素内酯调控植物耐盐机理研究进展[J]. 中国农业科技导报,2024,26(2):1-12.
[67]刘金隆,高会玲,洪立洲,等. 植物色素在24-表油菜素内酯调节油菜耐盐性中的作用[J]. 西北植物学报,2013,33(1):90-100.
[68]辛正琦,何会流. 外源EBR对NaCl胁迫下颠茄幼苗光合色素含量及荧光特性的调节[J]. 内江师范学院学报,2021,36(12):55-60,75.
[69]AI Y, CHEN Y L, WANG N, et al. Overexpression of MtIPT gene enhanced drought tolerance and delayed leaf senescence of creeping bentgrass (Agrostis stolonifera L.)[J]. BMC Plant Biology,2024,24(1):734.
[70]TACHIBANA R, YAMAGAMI A, MIYAGI S, et al. BRZ-INSENSITIVE-PALE GREEN 1 is encoded by chlorophyll biosynthesis enzyme gene that functions in the downstream of brassinosteroid signaling[J]. Bioscience,Biotechnology,and Biochemistry,2022,86(8):1041-1048.
[71]LEE S H, KIM S H, PARK T K, et al. Transcription factors BZR1 and PAP1 cooperate to promote anthocyanin biosynthesis in Arabidopsis shoots[J]. The Plant Cell,2024,36(9):3654-3673.
[72]WANG Y C, ZHU Y S, JIANG H Y, et al. The regulatory module MdBZR1-MdCOL6 mediates brassinosteroid- and light-regulated anthocyanin synthesis in apple[J]. New Phytologist,2023,238(4):1516-1533.
[73]SANG K Q, LI J J, QIAN X J, et al. The APETALA2a/DWARF/BRASSINAZOLE-RESISTANT 1 module contributes to carotenoid synthesis in tomato fruits[J]. The Plant Journal,2022,112(5):1238-1251.
[74]ZHOU Y Z, CHENG Y X, ZHONG R, et al. Brassinolide and gibberellin promote grape fruit development and quality[J]. Scientia Horticulturae,2024,338:113619.
[75]高金山,边庆华,张永忠,等. 细胞分裂素CPPU的研究进展[J]. 农药,2006,45(3):151-154.
[76]ZHU L H, LIAO Y W, LIN K, et al. Cytokinin promotes anthocyanin biosynthesis via regulating sugar accumulation and MYB113 expression in Eucalyptus[J]. Tree Physiology,2024,44(1):tpad154.
[77]万华方,卢东,梁颖,等. 细胞分裂素对甘蓝型油菜种皮色泽形成的影响[J]. 作物学报,2012,38(10):1900-1907.
[78]DUAN L J, ZENG Z Y, TANG Y D, et al. New insights into the role of cytokinin in regulating anthocyanin biosynthesis and leaf expansion:an integrated transcriptomic,metabolomic,and physiological analysis of Hypericum monogynum[J]. Forests,2025,16(3):521.
[79]TAN C X, YANG J Y, XUE X Y, et al. MsMYB62-like as a negative regulator of anthocyanin biosynthesis in Malus spectabilis[J]. Plant Signaling & Behavior,2024,19(1):2318509.
[80]XIAO H, NIE R, ZENG Y, et al. The physiological mechanism of the flower color formation of Camellia reticulata ‘Tongzimian’[J]. Ornamental Plant Research,2025,5(1):1-12.
[81]罗秀芹,薛晶晶,蔡杰,等. GA3处理对木薯类胡萝卜素生物合成的影响[J]. 热带作物学报,2022,43(2):346-352.
[82]CHEN K Q, ZHAO X Y, AN X H, et al. MdHIR proteins repress anthocyanin accumulation by interacting with the MdJAZ2 protein to inhibit its degradation in apples[J]. Scientific Reports,2017,7:44484.
[83]MARN-DE LA ROSA N, PFEIFFER A, HILL K, et al. Genome wide binding site analysis reveals transcriptional coactivation of cytokinin-responsive genes by DELLA proteins[J]. PLoS Genetics,2015,11(7):e1005337.
[84]UM T, CHOI J, PARK T, et al. Rice microRNA171f/SCL6 module enhances drought tolerance by regulation of flavonoid biosynthesis genes[J]. Plant Direct,2022,6(1):e374.
[85]北京大学发现光与赤霉素协同调控植物发育新机制[J]. 食品工业,2016,37(9):57.
[86]XIONG H B, LU D D, LI Z Y, et al. The DELLA-ABI4-HY5 module integrates light and gibberellin signals to regulate hypocotyl elongation[J]. Plant Communications,2023,4(5):100597.
[87]ZHENG T, DONG T Y, HAIDER M S, et al. Brassinosteroid regulates 3-hydroxy-3-methylglutaryl CoA reductase to promote grape fruit development[J]. Journal of Agricultural and Food Chemistry,2020,68(43):11987-11996.
[88]CHEN C, WANG L X, LI M Y, et al. Effects of exogenous 24-epibrassinolide leaves spraying application on chlorophyll accumulation and gene expression profiles of chlorophyll metabolism in celery[J]. Horticulturae,2023,9(12):1285.
[89]SONG R F, HU X Y, LIU W C, et al. ABA functions in low phosphate-induced anthocyanin accumulation through the transcription factor ABI5 in Arabidopsis[J]. Plant Cell Reports,2024,43(2):55.
[90]WAN Q Q, LU M, JIANG G G, et al. The characterization of OfRGA in regulation of flower size through tuning cell expansion genes[J]. Frontiers in Plant Science,2024,15:1502347.
[91]ZHANG J H, LI L J, ZHANG Z W, et al. The effect of ethephon on ethylene and chlorophyll in Zoysia japonica leaves[J]. International Journal of Molecular Sciences,2024,25(3):1663.
[92]FAN Z Q, TAN X L, CHEN J W, et al. BrNAC055,a novel transcriptional activator,regulates leaf senescence in Chinese flowering cabbage by modulating reactive oxygen species production and chlorophyll degradation[J]. Journal of Agricultural and Food Chemistry,2018,66(36):9399-9408.
[93]LI J, LI J, LUO J H, et al. Brassinosteroid-mediated carotenoid regulation enhances chilling tolerance in pepper[J]. BMC Genomics,2025,26(1):601.
[94]AN J P, ZHANG X W, BI S Q, et al. The ERF transcription factor MdERF38 promotes drought stress-induced anthocyanin biosynthesis in apple[J]. The Plant Journal,2020,101(3):573-589.
[95]HAN T Y, WU W L, LI W L. Transcriptome analysis revealed the mechanism by which exogenous ABA increases anthocyanins in blueberry fruit during veraison[J]. Frontiers in Plant Science,2021,12:758215.

相似文献/References:

[1]刘强,柳正葳,刘召亮,等.一氧化氮对铝胁迫下烟草根系营养元素吸收和呼吸电子传递及内源激素含量的影响[J].江苏农业学报,2020,(02):292.[doi:doi:10.3969/j.issn.1000-4440.2020.02.006]
　LIU Qiang,LIU Zheng-wei,LIU Zhao-liang,et al.Effects of nitric oxide on nutrient uptake, respiratory electron transfer and endogenous phytohormones contents in roots of tobacco seedlings under aluminum stress[J].,2020,(06):292.[doi:doi:10.3969/j.issn.1000-4440.2020.02.006]
[2]戴毅,田龙果,潘贞志,等.激素和非生物逆境胁迫调控植物硝酸盐转运蛋白功能的研究进展[J].江苏农业学报,2020,(06):1595.[doi:doi:10.3969/j.issn.1000-4440.2020.06.033]
　DAI Yi,TIAN Long-guo,PAN Zhen-zhi,et al.Research progress on the regulation of plant nitrate transporter functions by hormones and abiotic stress[J].,2020,(06):1595.[doi:doi:10.3969/j.issn.1000-4440.2020.06.033]
[3]王江英,葛金涛,邵小斌,等.筛选白花油茶胚性愈伤组织诱导成苗的关键激素通路及相关基因[J].江苏农业学报,2025,(03):599.[doi:doi:10.3969/j.issn.1000-4440.2025.03.019]
　WANG Jiangying,GE Jintao,SHAO Xiaobin,et al.Screening of key hormone pathways and related genes for the induction of plantlets from embryogenic callus of Camellia oleifera[J].,2025,(06):599.[doi:doi:10.3969/j.issn.1000-4440.2025.03.019]

备注/Memo

备注/Memo:
收稿日期：2025-09-24基金项目：广东省林业创新项目(2021KJCX002)；广东省基础与应用基础研究基金项目(2023A1515140021)作者简介：郑绿珠(1999-),女,广东普宁人,硕士研究生,研究方向为植物生理与分子生物学。(E-mail)lvzhuzheng@163.com通讯作者：蔡志全,(E-mail)zhiquan.cai@126.com
更新日期/Last Update: 2026-07-15